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HARVARD UNIVERSITY

LIBRARY

OF THE

Museum of Comparative Zoology

GENERAL CATALOGUE

+7/ 7

OF THE

HOMOPTERA

Fascicle VII CERCOPOIDEA Part 3

APHROPHORIDAE

ins. m*. zoiit

FE B 2 1 1962

harvard

bmsjh

BY

Z. P. METCALF, D.Sc.

State College of the University of North Carolina

Contribution from the Entomology Department, North Carolina Agricultural Experiment Station, Raleigh, C., U. S. A. Published, by North Carolina State College, Raleigh, N. C. with the approval of the Director of Research, as Paper No. 1324.

Printed by a Grant in Aid from the National Science Foundation

1962

GENERAL CATALOGUE

OF THE

HOMOPTERA

Fascicle VII CERCOPOIDEA Part 3

APHROPHORIDAE

BY

Z. P. METCALF, D.Sc.

State College of the University of North Carolina

I

Contribution from the Entomology Department, North Carolina Agricultural Experiment Station, Raleigh, N. C., U. S. A. Published by North Carolina State College, Raleigh, N. C. with the approval of the Director of Research, as Paper No. 1324.

Printed by a Grant in Aid from the National Science Foundation

1962

Library of Congress Catalog Card Number 28-7846

MUS. COMP. 2031 LIBRARY

FEB 2 1 1962

HARVARD

UNIVERSITY

Printed in the United States of America by WAVERLY PRESS, INC., Baltimore, Md.

INTRODUCTION

A catalogue of animals should serve practically all fields of biology. It should be, therefore, as complete a listing of all the records of families, subfamilies, tribes, genera, species, and varieties as it is possible for the author to assemble. The nature of the reference is of the greatest impor¬ tance. A student of zoogeography should be able to find a complete list of the regions inhabited by the various species. The student of ecology should be able to find references to all that is known about the life history, food plants, and other pertinent data. The student of economic entomology should be able to check the histories of those species that are suspected of being injurious to plants. The plant disease specialist should be able to check the current nomenclature of the species which are disease vectors or suspected of being disease vectors. Those research workers studying morphology, physiology, or genetics should be able to trace the develop¬ ment in the field of their special interest. And lastly, the student of tax¬ onomy should be relieved of the burden of searching for past recordings, and the journals which publish taxonomic papers should not have to pay the cost of publishing the past records and duplicating the synonymy that is already well known. Thus it would be necessary only to record synonymy which has been developed since the publication of the catalogue.

In this as in other catalogues of the present series the family is divided into genera which are arranged in as nearly a phylogenetic order as our present knowledge will permit. The species are arranged under the genera in alphabetic sequence.

The notes which follow the references are generally self-explanatory, but three points may be mentioned here. Such notations as “[described],” “[notes],” “[key],” and the like are intended to be suggestive rather than precise or exclusive. The notation “[error]” means not accepted in this catalogue. Usually the latest published synonymy is accepted, but not always. The notation “[comparative note]” is used to designate those references, often of the greatest taxonomic significance, in which two generic, specific, or other groups are compared. All references have been checked against the original save those marked with an asterisk (*), which have been accepted from reliable sources. Every effort has been made to have the references full and complete and to give an indication of the character of the data contained. Where the writer knows that reprints have been issued with different pagination, this is indicated in parentheses fol¬ lowing the page numbers of the original. In a few cases, where the matter has been reprinted under a different title with different pagination, separate references are given. Where authentic editions have been republished these

in

IV

General Catalogue of the Homoptera

have been indicated by separate references. Genera established without included species have been dated from their original description, with their type fixation dated at the earliest time the action can logically be inter¬ preted to have occurred. Original spellings have been retained in all cases, except obvious typographical errors. Typographical errors are indicated by placing “[sic]” after the misspelled name. This is not for the purpose of calling attention to the mistakes authors have made but to make certain that students following the catalogue have a clear understanding of what is implied.

The determination of obvious typographical errors is not always easy. These errors range all the way from clearly apparent transpositions of letters, to misspelled names of individuals, geographical names, and many others of like character. However, so many generic names have been formed by the transposition and substitution of letters that one is often at a loss to know which names are misspelled and which are not. For example, Neokolla and Keonolla, or Alebra , Elabra, and Eualebra.

All variations in spelling, save those which refer to endings only, are ap¬ propriately cross-referred. The species have also been listed under every known generic combination so that it is possible to find any species with¬ out recourse to the index, which will be published at the completion of the Fascicle.

The writer takes no pride in the authorship of nomina nova. He uses them only when he thinks they are necessary. No new names are proposed for species or genera described by living authors without first giving the original author an opportunity to correct primary or secondary homonyms.

A catalogue is not a revision of a group and the location of the genera and species in the present catalogue is merely an attempt to reflect what is believed to be the general consensus of the best authorities at the present time. Doubtful genera and species have been included to call them to the attention of subsequent revisers.

An attempt has been made throughout this catalogue to interpret all references in the light of the period in which they were published. This is not always easy and leads perhaps to erroneous conclusions at times, but it is certainly much sounder from the standpoint of nomenclature than the tendency to evaluate references from previous decades in the light of present day practices and procedures.

In this series we have departed from our previous method of giving latinized names to varieties which in the past have been designated by Arabic letters, Greek letters, numbers, or other designations. All varieties are catalogued, however, under the first designation regardless of its form. Some of these varieties may prove to be valid species with further study and then may be given latinized designations.

Cercopoidea Aphrophoridae

V

Primary homonyms of species have been replaced by nomina nova ; secondary homonyms have been restored to their original names if the homonymy has been relieved by moving one or the other or both species to other genera.

All specific endings have been modified to agree with the suggestions made by Blackwelder 1940, Journal of the Washington Academy of Sci¬ ences 31: 135-140. There are certain disadvantages inherent in this method, but it seems to the writer that these limitations are outweighed by the advantages of this system. We are confronted by so many coined words that can have no sex that it seems better to use this system and disregard apparent violations of Latin and Greek grammar. Where the sex of generic names can be determined the appropriate adjustments are made in the specific names.

In a detailed catalogue such as this it is practically impossible to elimi¬ nate all errors. We will appreciate it very much if our attention is directed to all such mistakes. It is especially easy to overlook local lists when pub¬ lished in the proceedings of local natural history societies, references to species of economic importance when published in scientific agricultural journals, references to vectors of plant diseases when published in phyto- pathological journals, and many other references of a similar nature. It will be very much appreciated if authors will send us reprints of all articles dealing with the Homoptera. Also it would seem that changes in the nomenclature adopted in this catalogue should not be made in the future without a full explanation.

All literature available to the writer before December 31, 1955, has been included in this catalogue.

The present catalogue of the Family Aphrophoridae is the third part of the Cercopoidea. This was virtually complete at the time of the death of Dr. Z. P. Metcalf on January 5, 1956. It was completed with the aid of Miss Virginia Wade, Research Analyst, N. C. State College, who worked with Dr. Metcalf for seven years, and the technical advice of Dr. David A. Young, N. C. State College, and Dr. W. H. Anderson, Insect Identifica¬ tion and Parasite Introduction Laboratories, U. S. Department of Agri¬ culture, Beltsville, Maryland. The higher categories utilized in this classi¬ fication are those decided upon by Dr. Metcalf during his work on the catalogue.

In this part of the catalogue a method of literature citation has been adopted which differs from that used in the catalogues of the fulgoroid families. This change has been instituted to save time, space, and publica¬ tion costs. It is believed that the revised method of literature citation will cause no great inconvenience to the reader. Dr. Metcalf himself advocated it under certain conditions in the past, and it is felt that the use of abbrevi-

VI

General Catalogue of the Homoptera

ated literature citations in his catalogues would have met with his ap¬ proval.

Under the abbreviated system used here, and to be used for subsequent portions of the Cercopoidea catalogue, the literature references are cited by author, year, and key letter, e.g. Fabricius 1802a. The full reference can be obtained by consulting the Bibliography of the Cercopoidea in which authors are arranged alphabetically and their works chronologically under the author’s name. This method of citation eliminates the necessity for a complete literature citation in the catalogue proper.

The Family Aphrophoridae consists of 151 genera and 820 species. Four invalid genera are included. This family may be characterized as follows: Eyes with horizontal diameter greater than vertical diameter; pronotum with anterior margin strongly convex or subangulate, anterolateral mar¬ gins usually short; scutellum flat, not spinous, shorter than pronotum; fore wing with clavus acute or obliquely truncate at apex.

The following classification has been adopted for this family:

Page

Family Aphrophoridae . 1

Subfamily Neaninae . 9

Subfamily Aphrophorinae . 10

Tribe Ptyelini . 10

Tribe Takagini . 90

Tribe Cloviini . 91

Tribe Orthoraphini . 151

Tribe Lepyroniini . 153

Tribe Philaenini . 222

Tribe Aphrophorini . 430

Tribe Philagrini . 571

Twenty-five of the species in the present catalogue have a distribution covering two or more zoogeographic regions. The localities for four species are unknown. The other species are recorded as follows: 297 from the Palearctic Region, 103 from the Ethiopian Region, 100 from the Malaysian Region, 89 from the Austromalayan Region, 68 from the Oriental Region, 43 from the Neotropical Region, 35 from the Caribbean Region, 20 from the Oceanic Region, 19 from the Nearctic Region, 16 from the Australian Region, and one from the Maorian Region.

The known geographic distribution of each species is indicated by super¬ script figures at the end of the lines, which correspond to geographic re¬ gions designated by the same superscript figures. In general the distribu¬ tion is as given by the author of the reference, using the country as the smallest unit except in the larger countries where states and provinces are

Cercopoidea Aphrophoridae

vii

used as the smallest units. In the larger island groups the individual islands are indicated wherever possible in view of the importance of island en¬ demism.

Z. P. Metcalf and

D. A. Young

Raleigh , North Carolina May 1961

Family APHROPHORIDAE Amyot and Serville

Aphrophorides

A phrophoroideae Aphrophorides

Aphrophoridae A phrophoridum Aphrophoridae

Aphrophorides

Aphrophoridae

Aphrophorida

Aphrophorina

Aphrophoridae

Aphrophorides

Aphrophoridae

Aphrophorides

Aphrophoridae

Aphrophorina

Aphrophorinae

Aphrophoridae

Amyot and Serville 1843a: 563 Group [Described] Agassiz, Erichson, and Germar 1846a: 2 [Listed] Agassiz 1848a: 81 [Listed]

Spinola 1850a: 56 Subfamily [Key]

Walker 1851b: 697 Subtribe [New species from various regions]

Walker 1857a: 96 Subtribe [Listed]

Walker 1857b: 166 Subtribe [Listed]

Walker 1858a: 93 Subtribe [Listed]

Dohrn 1859a: 68 [Listed]

Stal 1859b: 286 Family [Listed]

Walker 1860a: 18 [List of genera and species from Great Britain]

Favarcq 1861a: 80 Group [Listed]

Fuss 1862a: 17 Family [List of species from Tran¬ sylvania]

Morris 1865a: 111 [Listed]

Stal 1866a: 55 Subfamily [Key] :66 [Key to genera] :262 [Listed]

Dallas 1867a: 569 Subfamily [Key]

Stal 1869a: 15 Subfamily [Listed]

Stal 1870c: 722 Subfamily [Review of species from the Philippine Islands]

Walker 1870c: 2404 Family [Listed]

Sahlberg 1871a: 66 Tribe [Key to genera from Europe] :82 [Described]

Scott 1871d: 241 Family [Described, key to genera from Great Britain] Equals Aphrophorida Provancher 1872b: 351 [New species from Canada] Butler 1874f : 26 Family Equals Aphrophorides Douglas and Scott 1876a: 71 [Listed]

Bloomfield 1878a: 38 [Listed]

Berg 1879c: 219 Subfamily Equals Aphrophorida Berg 1879e: 238 Subfamily Equals Aphrophorida Distant 1879e: 14 Subfamily Equals Aphrophorina White 1879a: 214 Subfamily [List of species from New Zealand]

Cooke 1882a: 71 [List of species from England] Bloomfield 1883a: 34 [List of species from England]

1

2

General Catalogue of the Homoptera

Aphrophorina

Aphrophorida

Aphrophorina

Aphrophorida

Aphrophorinae

Aphrophorida

Aphrophorideen

Aphrophoriden

Aphrophorida

Aphrophorinae

Aphrophorina

Aphrophorinae

Aphrophoridae

Aphrophorinae

Aphrophoridae

Aphrophorinae

Aphrophorida

Aphrophorina

Berg 1884b: 116 Subfamily [Listed]

Berg 1884e: 199 Subfamily [Listed]

Uhler 1884a: 243 Subfamily [Notes]

Atkinson 1885a: 18 Subfamily [Described] Equals Aphrophorida

Uhler 1888c: 243 Subfamily [Notes]

Hansen 1890a: 28 [Listed]

Hansen 1890a : 43 [Morphology of wings]

Smith 1890a: 442 Subfamily [Listed]

Distant 1892a: 248 Subfamily [Listed]

South wick 1892b: 318 Subfamily [Listed]

Van Duzee 1892d: 296 Subfamily [Listed]

Karsch 1894a: 114 [Listed]

Karsch 1894a: 114 [List of species from Cameroons] Van Duzee 1894e: 193 Subfamily [List of genera and species from New York]

Goding 1895b: 483 Subfamily [Key] :484 [Key to genera from North America]

Uhler 1896a: 285 Subfamily [New species from Japan]

Ball 1899a: 205 [Synonymy] :207 Subfamily [Key] :209 [Notes, key to genera]

Haglund 1899a: 52 Subfamily [Review of species from Cameroons]

Hansen 1900b: 171 [Morphology of wings]

Kirby 1900a: 130, 137 [Listed]

Smith 1900a: 90 Subfamily [Listed]

Matsumura 1903a: 17 Subfamily [Key] :20 [De¬ scribed, key to genera from Japan]

Ashmead 1904a: 131 Subfamily [List of species from Alaska]

Hutton 1904a: 224 Subfamily [Catalogue of species from New Zealand]

Jacobi 1904a: 775 Subfamily [New species from Northeastern Africa]

Matsumura 1904a: 33 Subfamily [Listed]

Wirtner 1904a: 216 Subfamily [List of genera and species from Pennsylvania]

Kirkaldy 1906c: 290 [Key]

Oshanin 1906a: 25 Subfamily [Catalogue of genera and species from the Palearctic Region] Equals Aphrophorides Equals Aphrophorida

Cercopoidea Aphrophoridae

3

Aphrophorinae

Aphrophorine

Aphrophorinae

Aphrophoridae

Aphrophorinae

Aphrophorina

Aphrophorinae

Aphrophoridae

Aphrophorinae

Matsumura 1907b: 107 Subfamily [Listed]

Distant 1908c: 310 Subfamily [Notes on genera and species from Ethiopia]

Distant 1908g: 79 Subfamily [Key] :85 [Described, catalogued, key to genera] Equals Aphro- phorides Equals Aphrophorida Equals Aphro¬ phorina Equals Ptyelinae Equals Cercopinae [in part.]

Distant 1908h: 219 Subfamily [Catalogued] Equals Cercopinae [in part.] Equals Aphrophorides Equals Aphrophorida Equals Aphrophorina Equals Ptyelinae

Distant 1909a: 188 Subfamily [Notes on genera and species from the Neotropical Region]

Distant 1909h: 45 Subfamily [Listed]

Distant 1909i: 81 Subfamily [Listed]

Distant 1909j : 174 Subfamily [Listed]

Lefroy and Howlett 1909a: 732 [Notes] :733 [De¬ scribed]

Banks 1910a: 49 Subfamily [Listed]

Jacobi 1910b: 113 Subfamily [New species from Tanganyika]

Lefroy 1910a: 235 Division [Listed]

Schmidt 1910g: 337 Subfamily [Listed]

Shiraki 1910a: [79] [Listed]

Smith 1910a: 98 Subfamily [Listed]

Matsumura 1911b: 18 [List of species from Sak¬ halin]

Schmidt 1911b: 287 Subfamily [Listed]

Haupt 1912b: 448 (3) Subfamily [List of species from Thuringia]

Lallemand 1912a: § Subfamily [Key] :18 [Described, catalogued, key to genera] Equals Aphro¬ phorides Equals Aphrophorida Equals Aphro¬ phorina Equals Ptyelinae Equals Cercopinae [in part.]

Licent 1912a: 77 [Physiology of digestive system] Oshanin 1912a: 97 [List of genera and species from the Palearctic Region]

Schmidt 1912e: 170 Subfamily [Listed]

Schmidt 1912g: 150 Subfamily [Listed]

Schumacher 1912a: 322 Subfamily [Listed]

4

General Catalogue of the Homoptera

Aphrophoridae

Aphrophorinae

Aphrophorida

Aphrophorinae

Aphrophinae [sic] Aphrophorinae

Shiraki 1913a: [154] [Listed]

Distant 19141: 389 Subfamily [Listed]

Maki 1914a: 354 [Listed]

Melichar 1914b: 116 Subfamily [New species from Java]

Van Duzee 1914a: 48 Family [List of species from California]

Distant 1915d: 506 Subfamily [Listed]

Matsumura 1915a: 157, 183 [List of species from Korea]

Metcalf 1915a: 47 (15) Subfamily [List of species from North Carolina]

Schumacher 1915a: 87 Subfamily [Listed] Schumacher 1915b: 113 [Listed]

Cogan 1916a: 169 Subfamily [Key] :173 [Key to genera from South Africa]

Distant 1916a: 185 Subfamily [Listed]

Matsumura 1916b: 414 [Key, notes, key to genera] Oshanin 1916a: 74 Subfamily [Bibliography]

Van Duzee 1916a: 57 Subfamily [List of genera and species from the Nearctic Region] Equals Ptyelinae Equals Cercopinae [in part.] Deletang 1917a: 263 Subfamily [Key]

Deletang 1917a: 264 Subfamily [Notes, key to genera from Argentina] Equals Ptyelinae Equals Aphrophorides Equals Cercopinae [in part.] Equals Aphrophorina Equals Aphro¬ phorida

Distant 1917a: 306 Subfamily [Listed]

Jacobi 1917a: 544 Subfamily [New species from Madagascar]

Matsumura 1917b: 414 [Key, notes, key to genera] Metcalf 1917a: 28 Subfamily [Tracheation]

Van Duzee 1917b: 509 Subfamily [Catalogued] Equals Aphrophoridae Equals Aphrophorida Equals Aphrophorina Equals Cercopinae [in part.] Equals Ptyelinae Baker 1919a: 67 [Notes]

Haupt 1919a: 157 Subfamily [Described] :161 [Key to genera from Europe] :162 [Notes]

Schmidt 1919f: 379 Subfamily [Listed]

Cercopoidea Aphrophoridae

5

Aphrohorinae [sic] Aphrophorinae

Distant 1920f: 463 Subfamily [New species from New Caledonia]

Lallemand 1920a: 260 [New species from Africa] Lallemand 1920d: 280 Subfamily [New species from Madagascar]

Matsumura 1920a: 345 [Review of species from Japan]

Moore 1920a: 23 Subfamily [List of species from Canada]

Schmidt 1920a: 48 Subfamily [Listed]

Schmidt 1920k: 115 Subfamily [Listed]

Jacobi 1921a: 4 Subfamily [Notes]

Doering 1922a: 515 Subfamily [Described]

Haupt 1922a: 17 Subfamily [Described] :19 [Key to genera from Europe] :20 [Listed]

[Kazui] 1922a: 193 [Listed]

Lallemand 1922a: 273 [New species from the Philippine Islands]

Lallemand 1922b: 64 Subfamily [Listed]

Schmidt 1922a: 1 Subfamily [Listed]

Schmidt 1922c: 176 Subfamily [Listed]

Schmidt 1922d: 180 Subfamily [Listed]

Schmidt 1922e: 183 Subfamily [Listed]

Deletang 1923a: 584 [Key, notes, key to genera from Argentina]

Goding 1923a: 1 [Key] :2 Subfamily [Key to genera from Ecuador] :7 [Listed]

Aphrophinae [sic] Aphrophorinae

Stearns 1923a: 209 [Key]

Stearns 1923a: 211 Subfamily [Described, key to genera]

Lallemand 1924b: 202 Subfamily [Listed]

Buchner 1925a: 90 Subfamily [Listed] :102-121 [Symbionts]

China 1925c: 481 Subfamily [Listed]

Handlirsch 1925a: 1114 Subfamily [Described] Equals Aphrophorides Equals Aphrophoridae Equals Aphrophorida Equals Aphrophorina Equals Ptyelinae Equals Cercopinae [in part.] Equals Clastopteridae [in part.]

Hesse 1925a: 173 Subfamily [List of genera and species from southwestern Africa]

Kupka 1925a: 104 Subfamily [Listed]

6

General Catalogue of the Homoptera

Singh-Pruthi 1925b: 200 Subfamily [Listed] :202 [Comparative notes, notes on d' genitalia] Kuwayama 1926a: 20 Subfamily [Listed]

Schmidt 1926a: 224 Subfamily [Listed]

Downes 1927a: 16 Subfamily [List of species from British Columbia]

Lallemand 1927b: 101 Subfamily [New species from various regions]

Dozier 1928a: 8 [Listed]

Lallemand 1928c: 48 [New species from Samoa] Lallemand 1928e: 197 Subfamily [List of species from Western Sumatra]

Leonard 1928a: 145 Subfamily [List of genera and species from New York]

Schmidt 1928b: 112 Subfamily [Listed]

Box 1929a: 12 Subfamily [Notes]

Haupt 1929c: 221 Subfamily [Key] :222 [Key to genera]

Kuznezov 1929a: 308 Subfamily [List of species from Turkestan]

Kuznezov 1929f: 32 [List of species from Turkestan] Lallemand 1929c: 86 Subfamily [List of species from Belgian Congo]

Luigioni 1929a: 217 (19) Subfamily [Listed]

Doering 1930a: 54 Subfamily [Key]

Pendlebury 1930a: 108 Subfamily [Listed]

Schmidt 1930b: 113 Subfamily [Listed] de Seabra 1930a: 317 Subfamily [Listed] de Seabra 1930b: 241 Subfamily [Listed]

Aphrophoridae

Kato 1931c: 60 [Listed]

Kato 1931d: 163 (7) Family [A new species from Formosa]

Aphrophorinae

Lallemand 1931d: 78 Subfamily [New species from the East Indies]

Aphrophorina

Marelli 1931a: 14 Subfamily [Listed]

Aphrophorinae

Schmidt 1931b: 66 Subfamily [Listed]

Dover 1932c: 35 [Listed]

Aphrophoridae

Kato 1932a: 139 Family [Listed]

Kato 1932b: 218 Family [Listed]

Aphrophorinae

Monte 1932a: 1 5 Subfamily [Listed]

Matsumura 1932b: 293 [Listed]

Schmidt 1932b: 55 Subfamily [Listed]

Cercopoidea Aphrophoridae

7

Aphrophoridae

Aphrophorinae

Aphrophoridae A phrophorinae

Aphrophoridae

Aphrophorinae

Aphrophoridae A phrophorinae

Aphrophoridae A phrophorinae

Aphrophoridae A phi ophorinae

Schmidt 1932c: 53 Subfamily [Listed]

Kato 1933c: 3 [Listed]

Kato 1933d: 14 [Listed]

Kato 1933q: 220 Family [New species from Japan] Lallemand 1933a: 2 [Listed]

Nast 1933a: 11 (5) [Taxonomy, notes]

Okamoto 1934a: 403 [Listed]

Silvestri 1934a: 328 Subfamily [Listed] :329 [Key to genera from Europe]

Haupt 1935a: 154 Subfamily [Key] :1 55 [Key to genera from Central Europe]

Wu 1935a: 34 Subfamily [Catalogued] Equals Aphrophorides Equals Aphrophorida Equals Aphrophorina Equals Plyelinae Equals Cerco- pinae [in part.]

Haku 1937a: 696 (22) [List of species from Korea] Lallemand 1938c: 3 Subfamily [Listed]

Matsumura 1938a: 147 Subfamily [Listed]

Nagaoka 1938a: 27 [List of species from Korea] Spooner 1938a: 25; Text fig. 1 [Phylogeny, illus¬ trated]

Lallemand 1939a: 3 Subfamily [Listed]

Lallemand 1939b: 54 Subfamily [Listed]

Lallemand 1939d: 57 Subfamily [Listed]

Lallemand 1939e: 101 Subfamily [Listed] de Seabra 1939c: 169 Subfamily [List of species from Portugal]

Evans 1940b: 70, 74, 75 Family [Notes]

Lallemand 1940b: 137 Subfamily [New species from Netherlands Indies]

Matsumura 1940a: 36 Subfamily [Key, key to genera] :67 [Notes] :75 [Listed]

Matsumura 1940b: 47 Subfamily [List of species from Manchuria]

Matsumura 1940d: 35 Subfamily [List of species from Formosa]

Nagaoka 1940a: 479 (47) [List of species from Korea]

Lallemand 1941a: 3, 4 [Listed] :5 Subfamily [List of species from Belgian Congo]

Evans 1942b: 39 [Venation, phylogeny]

Hayward 1942a: 36 Subfamily [Listed]

8

General Catalogue of the Homoptera

Aphrophoridae

Aphrophorinae

Aphrophoridae

A'phro'phorinae

[Aphrophoridae]

Aphrophorinae

Lallemand 1942b: 1 Subfamily [List of species from Belgian Congo]

Lallemand 1942d: 64 Subfamily [New species from Cameroons]

Lallemand 1942f: 69, 71 Subfamily [Listed]

Matsumura 1942b: 101 [Key to subfamilies] :102 [Key to genera]

de Seabra 1942a: 7 Subfamily [Listed]

Blote 1943a: 63 Subfamily [Key] :64 [Key to genera] Jacobi 1944a: 25 Family [New species from Fukien] :57, 63 [Zoogeography]

Kloet and Hincks 1945a: 51 [List of species from Great Britain]

Froeschner 1946a: 31 [Key]

Maa 1947a: 2 [Listed]

Dammerman 1948a: 153 [Listed]

Lutz 1948a: 78 [Notes]

Zachvatkin 1948a: 168 [Listed] :169 [Key to species] Zimmerman 1948a: 1 Subfamily [Listed] :18 [Re¬ view of genera and species from the Territory of Hawaii]

Lallemand 1949a: 8 Subfamily [Notes]

Muller 1949a: 354, 355 [Symbionts]

China 1950a: 243 Subfamily [Listed]

Esaki and Ishihara 1950a: 44 Subfamily [List of species from Northern China]

Kramer 1950a: 61; Chart 6 [Phylogeny, illustrated] Lallemand 1950b: 95 Subfamily [Listed]

Pesson 1951a: 1502 Subfamily Equals Cercopinae [in part.] :1505 [Notes]

Lallemand and Synave 1952a: 11 Subfamily [Listed] Lallemand and Synave 1952b: 13 [Listed]

Lallemand and Synave 1952c: 105 Subfamily [A new variety and new species from Madagas¬ car]

Lallemand and Synave 1952e: 26 Subfamily [Listed] Villiers 1952a: 215 [Key] :216 Subfamily [Described, review of genera and species from Africa] Buchner 1953a: 603 [Symbionts]

Lallemand and Synave 1953a: 234 [Listed] Lallemand and Synave 1953b: 190 [Listed] :195 Subfamily [New species]

Cercopoidea Aphrophoridae

9

Aphrophoridae

Aphrophorinae

Aphrophoridae

Mathur 1953a: 146 Subfamily [List of genera and species from India]

Dlabola 1954a: 134 Subfamily [Key] :135 [Review of genera and species from Czechoslovakia, cata¬ logued]

Lindberg 1954a: 189 Subfamily [Listed]

Metcalf 1954b: 13 Family [Listed]

Lallemand and Synave 1955b: 1 Subfamily [Listed] Metcalf 1955a: 264 Family [Listed]

Subfamily NEAENINAE Metcalf

Neaeninae Metcalf 1939a: 247 [Listed]

Metcalf 1944b: 152 [Bibliography]

Genus Neaenus Fowler

Haplotype Neaenus varius [n. sp.] Fowler 1897b: 191 Type Neaenus varius Lallemand 1912a: 21

Neaenus Fowler 1897b: 191 [gen. n.] :190 [Key]

Waterhouse 1902a: 236 [Listed]

Lallemand 1912a: 18 [Key] :21 [Described, cata¬ logued]

China 1925c: 482 [Comparative note]

Schulze, Kiikenthal, and Heider 1932b: 2200 [Cata¬ logued]

Neave 1940a: 273 [Catalogued]

Metcalf and Bruner 1944a: 109 [Key] :111 [De¬ scribed]

a Fowler (var.) [varius]. See Neaenus varius var. a Fowler.

b Fowler (var.) [varius]. See Neaenus varius var. b Fowler,

c Fowler (var.) [varius]. See Neaenus varius var. c Fowler.

d Fowler (var.) [varius]. See Neaenus varius var. d Fowler.

luteosignatus Valdes Ragues

Tomaspis luteosignatus Valdes Ragues 1910a: 445 [n. sp.]1 Tomaspis luteosignata Valdes Ragues 1914a: 105 [Listed]1 Neaenus luteosignatus Metcalf and Bruner 1944a: 111; PL 58, Fig. 9; PI. 59, Fig. 1 [Described, illustrated]1 Equals Tomaspis luteosignata Vald.

Localities: 1 Cuba, varius Fowler

Neaenus varius Fowler 1897b: 191; PI. 12, Figs. 1, 2 [n. sp., illustrated]1 N[eaenus] varius Lallemand 1912a: 22 [Catalogued]1 Localities: 1 Mexico.

10

General Catalogue of the Homoptera

varius var. a Fowler

[Neaenus varius] var. a Fowler 1897b: 191 [n. var.]1 [Neaenus varius] var. a Lallemand 1912a: 22 [Catalogued]1 Localities: 1 Mexico.

varius var. b Fowler

[Neaenus varius] var. b Fowler 1897b: 191 [n. var.]1 [Neaenus varius] var. b Lallemand 1912a: 22 [Catalogued]1 Localities: 1 Mexico.

varius var. c Fowler

[Neaenus varius] var. c Fowler 1897b: 191 [n. var.]1 [Neaenus varius] var. c Lallemand 1912a: 22 [Catalogued]1 Localities: 1 Mexico.

varius var. d Fowler

[Neaenus varius] var. d Fowler 1897b: 191 [n. var.]1 Localities: 1 Mexico.

Subfamily APHROPHORINAE Licent

Aphrophorinae Licent 1912a: 134 [Ecology]

Aphrophorini Lallemand 1933a: 3 [Listed]

Aphrophorinae Matsumura 1942b: 102 Subfamily [Key]

Jacobi 1944a: 58 [Zoogeography] Metcalf 1955a: 264 Subfamily [Listed]

Ptyelinae

Ptyelini Ptielini [sic]

Ptyelinae

Ptyelini

Ptylini [sic]

Ptyelini

Ptyelinae

Tribe PTYELINI Fowler

Fowler 1897a: 174 [Key]

Fowler 1897b: 190 Subfamily [Notes, key to genera from Central America]

Schmidt 1920k: 115 Tribe [Listed]

Lallemand 1924c: 294 Tribe [New species from New Caledonia and Loyalty Islands]

Lallemand 1925a: 6 Subfamily [Listed]

Lallemand 1925a: 6 Tribe [Listed]

Lallemand 1933a: 2 [Listed]

Lallemand 1935a: 679 Tribe [Listed]

Lallemand 1937a: 252 Tribe [List of species from Oceania]

Matsumura 1940a: 36 [Key, key to genera] Lallemand 1946a: 190 Tribe [Listed]

Maa 1947a: 2 [Listed]

Cercopoidea Aphrophoridae

1 1

Ptyelinae

Ptyelini

Ptyelini

Lallemand and Synave 1952a: 11 Tribe [Listed] Lallemand and Synave 1952b: 13 [Listed] Metcalf 1952a: 228 Tribe [Listed]

Franco 1953a: 6 [Key]

Lallemand and Synave 1953a: 234 Tribe [Listed |

Genus Tremapterus Spinola

Haplotype Tremapterus dregei [n. sp.] Spinola 1850b: 119 Type Tremapterus dregei Lallemand 1912a: 26

Tremapterus

Spinola 1850b: 117 [gen. n.] Spinola 1850a: 56 [Key] Schaum 1851a: 269 [Described]

Stal 1866a: 262 [Listed]

Marschall 1873a: 384 [Catalogued]

Scudder 1882b: 322 [Listed]

Jacobi 1904a: 778 [Comparative note]

Jacobi 1910b: 118 [Notes]

Lallemand 1912a: 18 [Key] :26 [Described, cata¬ logued]

Schumacher 1912a: 323 [A new species from Cam- eroons]

Schulze, Kukenthal, and Heider 1938a: 3519 [Cata¬ logued]

Neave 1940b: 528 [Catalogued]

Lallemand 1941a: 11 [Comparative note]

dregei Spinola

Tremapterus dregei Spinola 1850b: 119 [n. sp.]1 Tr[emapterus\ dregei Schaum 1851a: 269 [Listed]2 [Tremapterus] dregei Stal 1866a: 262 [Listed]3 Tremapterus dregei Jacobi 1910b: 118 [Notes]

T[remapterus] dregrei [sic] Lallemand 1912a: 27 [Catalogued]3 Localities: 1 Cape of Good Hope. 2 Natal. 3 Africa. dregrei. Typographical error for Tremapterus dregei Spinola.

major Jacobi

Tremapterus major Jacobi 1910b: 118; PI. 1, Figs. 32, 32a [n. sp., il¬ lustrated]1 :98, 135 [Listed]1

T[remapterus] major Lallemand 1912a: 27; PI. 1, Fig. 4 [Illustrated, catalogued]1

T[remapterus] major Schumacher 1912a: 323 [Comparative note] Tremapterus major Lallemand 1941a: 11 [Comparative note] Localities: 1 Tanganyika.

12

General Catalogue of the Homoptera

occidentalis Schumacher

Tremapterus occidentalis Schumacher 1912a: 323 [n. sp.]1 Localities: 1 Cameroons.

Genus Hemiapterus Jacobi

Haplotype Hemiapterus decurtatus [n. sp.] Jacobi 1904a: 777 Type H[emiapterus] decurtatus Distant 1908c: 311 Type Hemiapterus decurtatus Lallemand 1912a: 27

Hemiapterus Jacobi 1904a: 777 [gen. n.] :764 [Listed]

Melichar 1905a: 292 [Comparative note]

Distant 1908c: 311 [New species from Western Africa]

Jacobi 1912a: 31 [Comparative note]

Lallemand 1912a: 19 [Key] :27 [Described, cata¬ logued]

Waterhouse 1912a: 125 [Listed]

Jacobi 1921a: 5 [Taxonomy]

Schulze, Kiikenthal, and Heider 1930a: 1501 [Cata¬ logued]

Neave 1939b: 601 [Catalogued]

decurtatus Jacobi

H[emiapterus] decurtatus Jacobi 1904a: 778; PI. 44, Fig. 16 [n. sp., il¬ lustrated]1 :764 [Listed]2

H[emiapterus ] decurtatus Melichar 1905a: 292 [Comparative note] H[emiapterus\ decurtatus Lallemand 1912a: 27; PI. 1, Fig. 6 [Illustrated, catalogued]3

Hemiapterus decurtatus Jacobi 1921a: 5 [Listed]1 Localities: 1 Tanganyika. 2 Northeastern Africa. 3 Ethiopia.

fasciatus Distant. See Eulepyroniella fasciata Distant. fuscus Melichar. See Sepullia murrayi Signoret.

murrayi var. viridicollis Melichar. See Sepullia murrayi var. viridicollis Melichar. variegatus Distant. See Eulepyroniella fasciata var. variegata Distant. viridicollis Melichar. See Sepullia murrayi var. viridicollis Melichar.

Genus Sepullia Stal

Logotype S[epullia\ murrayi Distant 1908c: 311 Type Sepullia murrayi Schmidt 1911b: 288 Type Sepullia murrayi Lallemand 1912a: 27 Type S[epullia] murrayi Hesse 1925a: 174

Sepullia Stal 1866a: 79 [gen. n.] :69 [Key]

Cercopoidea Aphrophoridae

13

Dallas 1867a: 571 [Notes, catalogued]

Marschall 1873a: 380 [Catalogued]

Scudder 1882b: 290 [Listed]

Distant 1908c: 311 [Catalogued]

Distant 1908h: 224 [Listed]

Schmidt 1911b: 288 [Catalogued]

Lallemand 1912a: 19 [Key] :27 [Described, cata¬ logued]

Hesse 1925a: 174 [Catalogued]

Singh-Pruthi 1925b: 201; PL XXII, Fig. 175 [<?

genitalia described, illustrated]

Lallemand 1935b: 82 [Comparative note]

Schulze, Kiikenthal and Heider 1936b: 3168 [Cata¬ logued]

Neave 1940b: 176 [Catalogued]

Metcalf 1944b: 152 [Bibliography]

Metcalf 1947a: 161 [Zoogeography]

Lallemand and Synave 1952b: 17 [Catalogued, catalogue of species from Belgian Congo and Uganda]

_ Distant (var.) [ murrayi ]. See Sepullia murrayi var. nigromaculata Schmidt.

_ Jacobi (var.) [ curtula ]. See Sepullia murrayi var. unicolor Schmidt.

a St&l (var.) [vidua] . See Sepullia funebris var. a St&l. b St&l (var.) [vidua]. See Sepullia funebris var. b St&l. blemus. Typographical error for Sepullia blenna St&l.

blenna Stal

A [cocephalus] blennus Stal 1855a: 98 [n. sp.]1 Acocephalus blennus Walker 1858b: 261 [Listed]1 [Acucephalus] blennus Dohrn 1859a: 84 [Listed]2

S[epullia] blenna Stal 1866a: 80 [Described]2 Equals Acocephalus blennus Stal

[Sepullia blenna] Dallas 1867a: 571 Equals Acocephalus blennus Stal Sepullia blemus [sic] Signoret 1879a: 88 (42) [Taxonomy, catalogued] Equals A [cocephalus] blemus [sic] Stal S[epullia] blenna Distant 1908h: 224 [Catalogued]2 S[epullia] blenna Lallemand 1912a: 28 [Catalogued] Equals Acocepha¬ lus [blennus] Stal Localities: 1 Natal. 2 Caffraria.

brunea. Typographical error for Sepullia brunnea Lallemand. See Patriziana brunnea Lallemand.

brunnea Lallemand. See Patriziana brunnea Lallemand.

14

General Catalogue of the Homoptera

bukamensis Lallemand

Sepullia bukamensis Lallemand 1920a: 265 [n. sp.]1 Localities: 1 Belgian Congo. c St&l (var.) [vidua]. See Sepullia funebris var. c St&l.

callosa Lallemand

Sepullia callosa Lallemand 1935b: 81 [n. sp.]1 Localities: 1 Italian Somaliland.

curtula Jacobi. See Sepullia murrayi var. viridicollis Melichar. curtula var. _ Jacobi. See Sepullia murrayi var. unicolor Schmidt.

funebris Stal

A [cocephalus] funebris Stal 1855a: 98 [n. sp.]1 A[cocephalus] viduus Stal 1855a: 98 [n. sp.]1 Acocephalus funebris Walker 1858b: 261 [Listed]1 Acocephalus viduus Walker 1858b: 261 [Listed]1 [Acucephalus] funebris Dohrn 1859a: 84 [Listed]2 [Acucephalus] viduus Dohrn 1859a: 84 [Listed]2

S[epullia] vidua Stal 1866a: 81 [Described, catalogued, comparative note]2

Acocephalus funebris Stal 1866a: 81 To [Sepullia vidua] var. b Acocephalus viduus Stal 1866a: 81 To [Sepullia vidua] var. c [Sepullia vidua] Dallas 1867a: 571 Equals [Acocephalus] viduus Stal Sepullia vidua Signoret 1879a: 88 Equals Acocephalus viduus Stal Equals A[cocephalus] funebris Stal Sepullia vidua Schouteden 1901a: 122 [Listed]2 S[epullia] vidua Distant 1908h: 224 [Catalogued]2 S[epullia] vidua Lallemand 1912a: 28 [Catalogued]

Localities: 1 Natal. 2 Caffraria.

funebris var. a Stal

[Sepullia vidua] var. a Stal 1866a: 81 [n. var.]1 Localities: 1 Caffraria.

funebris var. b Stal

[Sepullia vidua] var. b Stal 1866a: 81 [n. var.]1 Equals Acocephalus funebris Stal Localities: 1 Caffraria.

funebris var. c Stal

[Sepullia vidua] var. c Stal 1866a: 81 [n. var.]1 Equals Acocephalus viduus Stal Localities: 1 Caffraria.

fuscolimbata Lallemand (var.). See Sepullia murrayi var. fuscolimbata Lallemand. fuseolimbata. Typographical error for Sepullia murrayi var. fuscolimbata Lallemand. impunctata Lallemand (var.). See Sepidlia murrayi var. impunctata Lallemand.

Cercopoidea Aphrophoridae

15

johannae Lallemand

Sepullia johannae Lallemand 1920a: 264 [n. sp.]1 Localities: 1 Belgian Congo.

murrayi Fairmaire and Signoret

Clastoptera murrayi Fairmaire and Signoret 1858a: 333 [n. sp.]1 Clastoptera murrayi Fairmaire and Signoret 1858b: 329 [n. sp.]1 S[epullia] murrayi Stal 1866a: 80 [Described]1 Equals Clastoptera murrayi Sign.

[Sepullia murrayi] Dallas 1867a: 571 Equals Clastoptera murrayi Sign. Hemiapterus fuscus Melichar 1905a: 292 [n. sp.]2 Sepullia murrayi Distant 1908c: 311 [Catalogued] Equals Clastoptera murrayi Sign.

Sepullia murrayi Schmidt 1911b: 288 [Notes, catalogued] Equals Clastoptera murrayi Sign. Equals Sepullia curtula Jac.

Sepullia murrayi Jacobi 1912a: 20 [Listed]3, 4 :31 [Notes, catalogued]5 Equals Hemiapterus viridicollis Mel. Equals Sepullia curtula Jac. S[epullia] murrayi Lallemand 1912a: 28 [Catalogued] Equals Hemiap¬ terus viridicollis Mel.

S[epullia] murrayi Lallemand 1920a: 265 [Comparative note]

Sepullia murrayi Jacobi 1921a: 5 [Catalogued] Equals H[emiapterus] fuscus Mel. Equals Hemiapterus viridicollis Mel.

Sepullia murrayi Lallemand 1929d: 213 [Catalogued] Equals Clastop¬ tera marrayi [sic] Sign. :214 [Key]

Sepullia murrayi Lallemand 1941a: 9 [Listed]5 Sepullia murrayi Lallemand and Synave 1954a: 909 [Listed]6 Localities: Malabar. 2 Tanganyika. 3 Tropical Western Africa. 4 Tropical Central Africa. 5 Belgian Congo. 6 Africa.

murrayi var. _ Distant. See Sepullia murrayi var. nigromaculata Schmidt.

murrayi var. curtula Jacobi. See Sepullia murrayi var. viridicollis Melichar.

murrayi var. fuscolimbata Lallemand

[Sepullia murrayi] fuscolimbata Lallemand 1929d: 213 [n. var.]1 Sepullia murrayi fuseolimbata [sic] Lallemand 1942b: 3 [Listed]1 Localities: 1 Belgian Congo.

murrayi var. fuseolimbata. Typographical error for Sepullia murrayi var. fuscolimbata Lallemand.

murrayi var. impunctata Lallemand

[Sepullia murrayi] impunctata Lallemand 1929d: 213 [n. var.]1 :214 [Key]

Sepullia murrayi impunctata Lallemand and Synave 1952b: 17 [Catalogued]1, 2

Localities: 1 Belgian Congo. 2 Uganda.

murrayi var. nigromaculata Schmidt

Sepullia murrayi var. _ Distant 1908c: 311 [n. var.]1

16

General Catalogue of the Homoptera

| Sepullia murrayi ] nigromaculata Schmidt 1911b: 289 Nom. nov. pro Sepullia murrayi var. _ Distant

[Sepullia murrayi] nigromaculata Lallemand 1929d: 214 [Catalogued]

Equals Sepullia murrayi var. _ Distant :215 [Key]

Sepullia murrayi nigromaculata Lallemand 1941a: 9 [Listed]2 Sepullia murrayi nigromaculata Lallemand 1942b: 2 [Listed]2 Localities: 1 Nigeria. 2 Belgian Congo.

murrayi var. transversa Lallemand

[Sepullia murrayi] transversa Lallemand 1929d: 213 [n. var.]1 :214 [Key] Localities: 1 Senegal.

murrayi var. unicolor Schmidt

[Sepullia curtula] var _ Jacobi 1910b: 118 [n. var.]1

[Sepullia murrayi] unicolor Schmidt 1911b: 289 Nom. nov. pro Sepullia curtula var. _ Jacobi

[Sepullia murrayi] unicolor Lallemand 1929d: 214 [Key, catalogued]

Equals Sepullia curtula var. _ Jac.

Sepullia murrayi unicolor Lallemand 1942b: 3 [Listed]2 Localities: 1 Tanganyika. 2 Belgian Congo.

murrayi var. viridicollis Melichar

Hemiapterus viridicollis Melichar 1905a: 291 [n. sp.]1 Sepullia curtula Jacobi 1910b: 118; PI. 1, Fig. 26 [n. sp., illustrated]1 :135 [Listed]1

Sepullia curtula Schmidt 1911b: 288 To Sepullia murrayi Sign. [Sepullia murrayi] curtula Schmidt 1911b: 289 [Notes]

Sepullia curtula Jacobi 1912a: 31 To Sepullia murrayi Sign. Hemiapterus viridicollis Jacobi 1912a: 31 To Sepullia murrayi Sign. Hemiapterus viridicollis Lallemand 1912a: 28 To S[epullia] murrayi Sign.

Hemiapterus viridicollis Jacobi 1921a: 5 To Sepullia murrayi Sign. [Sepullia murrayi] viridicollis Lallemand 1929d: 213 [Comparative note] :214 [Catalogued] Equals [Sepullia murrayi] curtula Schmidt Equals Sepullia curtula Jac. Equals Hemiapterus viridicollis Mel. :215 [Key]

Sepullia murrayi viridicollis Lallemand and Synave 1952b: 17 [Catalogued]2, 3 Equals Hemiapterus murrayi viridicollis Mel. Sepullia murrayi viridicollis Lallemand and Synave 1954a: 911 [Catalogued]4 Equals Hemiapterus [murrayi viridicollis Mel.] Localities: 1 Tanganyika. 2 Belgian Congo. 3 Uganda. 4 French Guinea.

nigromaculata Schmidt (var.). See Sepullia murrayi var. nigromaculata Schmidt.

nigropunctata Stal

S[epullia] nigropunctata Stal 1866a: 80 [n. sp.]1

Cercopoidea Aphrophoridae

17

Sepullia nigropunctata Dallas 1867a: 571 [Catalogued]2 S[epullia] nigropunctata Distant 1908h: 224 [Catalogued]1 S[epullia\ nigropunctata Lallemand 1912a: 28 [Catalogued]

Localities: 1 Cape